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By Peter Duren

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This "hybrid inviability" is one type of "post-zygotic" reproductive isolation. (iii) Gonadal barrier. " In his Notes on Hybridity(1862, 707) Darwin recognized that one barrier might occur first and then be replaced by another: "I suspect...

Having adopted the view that evolutionary change can only be brought about by natural selection, Darwin is confronted with a logical impasse. How can natural selection contribute to reproductive isolation when natural (phenotypic) selection is, after all, about selection of individuals for reproductive success, not for reproductive failure! However, Darwin continues: It may be admitted ... that it would profit an incipient species [variety] if it [the offspring] were rendered in some slight degree sterile when [members of the species were] crossed with ...

Darwin further notes (171): "An individual animal... " He is here contradicting himself in that he has already admitted (see above) that:"... it would profit an incipient species [variety] if it [its members] were rendered ... " This would indeed "indirectly give ... , the collective of individuals which constitutes an incipient species]. It will be shown later (chapter 4) that selective fertility (infertility with one group, while retaining fertility with another) might not necessarily result in the production of fewer offspring when viewed from Darwin's natural selection perspective.

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